Baxi K, Dorries K, Eisthen H. 2006. Is the vomeronasal system really specialized for detecting pheromones? Trends Neurosci. 29:1-7.
Bels VL, Chardon M, Kardong KV. 1994. Biomechanics of the hyolingual system in Squamata. In: Bels VL, Chardon M, Vandewalle P, editors. Advances in Comparative and Environmental Physiology vol. 18. Berlin: Springer. p. 197-240.
Bertmar G. 1981. Evolution of vomeronasal organs in vertebrates. Evolution. 35:359-366.
Burghardt GM. 1969. Comparative prey-attack studies in newborn snakes of the genus Thamnophis. Behaviour. 33:77-114.
Burghardt GM. 1970. Chemical perception in reptiles. In: Johnston JW, Moulton DG, Turk A, editors. Communication by chemical signals. New York: Appleton-Century-Crofts. p. 241-308.
Burghardt GM. 1993. The comperative imperative: genetics and ontogeny of chemoreceptive prey responses in natricine snakes. Brain Behav Evol. 41:138-146.
Burghardt GM, Pruitt CH. 1975. Role of the tongue and senses in feeding of naive and experienced garter snakes. Physiol Behav. 14:185-194.
Chiel HJ, Crago P, Mansour JM, Hathi K. 1992. Biomechanics of a muscular hydrostat: a model of lapping by a reptilian tongue. Biol Cybern. 67:403-415.
Cooper WE, Burghardt GM. 1990a. A comparative analysis of scoring methods for chemical discrimination of prey by squamate reptiles. J Chem Ecol. 16:45-65.
Cooper WE, Burghardt GM. 1990b. Vomerolfaction and vomodor. J Chem Ecol. 16:103-105.
Cowles RB, Phelan RL. 1958. Olfaction in Rattlesnakes. Copeia. 1958:77-83.
de Groot JH, van der Sluijs I, Snelderwaard PC, van Leeuwen JL. 2004. A three-dimensional kinematic analysis of tongue flicking in Python molurus. J Exp Biol. 207:827-839.
Gnanamuthu CP. 1937. Comparative study of the hyoid and tongue of some typical genera of reptiles. Proc Zool Soc Ser B. 107:1-63.
Goosse V, Bels VL. 1992. Kinematic analysis of tongue movements during chemosensory behavior in the European green lizard, Lacerta viridis (Reptilia, Lacertidae). Can J Zool. 70:1886-1896.
Gove D. 1979. A comparative study of snake and lizard tongue-flicking, with an evolutionary hypothesis. Z Tierpsych. 51:58-76.
Gove D, Burghardt GM. (1983). Context-correlated parameters of snake and lizard tongue-flicking. Anim Behav. 31:718-723.
Graves BM. 1993. Chemical delivery to the vomeronasal organs and functional domain of squamate chemoreception. Brain Behav Evol. 41:198-202.
Halpern M. 1976. The efferent connections of the olfactory bulb and accessory olfactory bulb in the snakes, Thamnophis sirtalis and Thamnophis radix. J Morphol. 150:553-578.
Halpern M. 1987. The organization and function of the vomeronasal system. Annu Rev Neurosci. 10:325-362.
Halpern M. 1992. Nasal chemical senses in reptiles: structure and function. In: Gans C, Crews D, editors. Biology of the Reptilia. Vol. 18. Chicago: University of Chicago. p. 424-532.
Halpern M, Kubie JL. 1980. Chemical access to the vomeronasal organs of garter snakes. Physiol Behav. 24:367-371.
Halpern M, Kubie JL. 1983. Snake tongue flicking behavior: clues to vomeronasal system functions. In: Silverstein RM, Müller-Schwarze D, editors. Chemical Signals III. New York: Plenum Publishing Corp. p. 45-72.
Halpern M, Martínez-Marcos A. 2003. Structure and function of the vomeronasal system: an update. Prog Neurobiol. 70:245-318.
Hansknecht KA. 2008. Lingual luring by mangrove saltmarsh snakes (Nerodia clarkii compressicauda). J Herpetol. 42:9-15.
Hansknecht KA, Burghardt GM. 2010. Stimulus control of lingual predatory luring and related foraging tactics of mangrove saltmarsh snakes (Nerodia clarkii compressicauda). J Comp Psychol. 124:159-165.
Hart BL, Leedy MG. 1987. Stimulus and hormonal determinants of flehmen behavior in cats. Horm Behav. 21:44-52.
Hartley RI, Sturm P. 1997. Triangulation. Comput Vis Image Und. 68:146-157.
Herrel A, Timmermans JP, De Vree F. 1998. Tongue flicking in agamid lizards: morphology, kinematics, and muscle activity patterns. Anat. Rec. 252:102-116.
Keiser ED. 1975. Observations on tongue extension of vine snakes (Genus Oxybelis) with suggested behavioral hypotheses. Herpetologica. 31:131-133.
Kier W, Smith K. 1985. Tongues, tentacles and trunks: the biomechanics of movement in muscular-hydrostats. Zool J Linn Soc Lond. 83:307-324.
Langebartel DA. 1968. The hyoid and its associated muscles in snakes. In: Bader RS, Heath JE, Selander RB, Smith HM, Wolfe RS, editors. Illinois biological monographs. Chicago: University of Illinois Press. p. 1-88.
Lanuza E, Halpern M. 1998. Efferents and centrifugal afferents of the main and accessory olfactory bulbs in the snake Thamnophis sirtalis. Brain Behav Evol. 51:1-22.
Leroy B, Toubeau G, Falmagne P, Wattiez R. 2006. Identification and characterization of new protein chemoattractants in the frog skin secretome. Mol Cell Prot. 5:2114-2123.
Martínez-Marcos A, Lanuza E, Halpern M. 2002. Neural substrates for processing chemosensory information in snakes. Brain Res Bull. 57:543-546.
Martínez-Marcos A, Ubeda-Bañón I, Halpern M. 2001. Neural substrates for tongue-flicking behavior in snakes. J Comp Neurol. 432:75-87.
Mason RT. 1992. Reptilian pheromones. In: Gans C, Crews D, editors. Biology of the Reptilia. Vol. 18. Chicago: University of Chicago. p. 114-228.
McDowell SB. 1972. The evolution of the tongue of snakes, and its bearing on snake origin. In: Dobzhansky T, Hecht MK, Steere WC, editors. Evolutionary biology. Vol. 6. New York: Appleton-Century-Crofts. p. 191-273.
Meredith M, Burghardt GM. 1978. Electrophysiological studies of the tongue and accessory olfactory bulb in garter snakes. Physiol Behav. 21:1001-1008.
Meredith M, Marques DM, O'Connell RO, Stern FL. 1980. Vomeronasal pump: significance for male hamster sexual behavior. Science. 207:1224-1226.
Mertens R. 1946. Die Warn- und drohreaktion der Reptilien. Abh Senck Naturforsch Ges. 471:1-10.
Mertens R. 1955. Die amphibien und Reptilien siidwestafrikas. Abh Senck Naturforsch Ges. 490:1-172.
Nowack C, Wöhrmann-Repenning A. 2009. New anatomical analyses suggest a pumping mechanism for the vomeronasal organ in anurans. Copeia. 2009:1-6.
Parsons TS. 1967. Evolution of the nasal structure in the lower tetrapods. Am Zool. 7:397-413.
Parsons TS. 1970. The nose and Jacobson's organ. In: Gans C, Parsons TS, editors. Biology of the Reptilia. Vol.2. New York: Academic Press. p. 99-191.
Pitman CRS. 1974: A guide to the snakes of Uganda. Codicote: Wheldon and Wesley.
Poran NS, Vandoros A, Halpern M. 1993. Nuzzling in the gray short-tailed opossum. I: Delivery of odors to vomeronasal organ. Physiol Behav. 53:959-967.
Schulterbrandt TG, Kubie J, von Gizycki H, Zuri I, Halpern M. 2008. Patterns of tongue-flicking by garter snakes (Thamnophis sirtalis) during presentation of chemicals under varying conditions. In: Hurst JL, Beynon RJ, Roberts JC, Wyatt TD, editors. Chemical signals in vertebrates XI. New York: Springer. p. 345-356.
Schwenk K. 1993. The evolution of chemoreception in squamate reptiles: a phylogenetic approach. Brain Behav Evol. 41:124-137.
Schwenk K. 1994. Why snakes have forked tongues. Science. 263:1573-1577.
Schwenk K. 1995. Of tongues and noses: chemoreception in lizards and snakes. Trends Ecol Evol. 10:7-12.
Schwenk K. 1996. Why snakes flick their tongues. Am Zool. 36:84A.
Seaman MA, Hill CC. 1996. Pairwise comparisons for proportions: a note on Cox and Key. Educ Psychol Meas. 56:452-459.
Sheffield LP, Law JH, Burghardt GM. 1968. On the nature of chemical food sign stimuli for newborn snakes. Comm Behav Biol Part A. 2:7-12.
Smargiassi M, Daghfous G, Leroy B, Legreneur P, Toubeau G, Bels V, Wattiez R. 2012. Chemical basis of prey recognition in thamnophiine snakes: the unexpected new roles of parvalbumins. PLoS One. 7:e39560.
Smith KK. 1984. The use of the tongue and hyoid apparatus during feeding in lizards (Ctenosaura similis and Tupinambis nigropunctatus). J Zool. 202:115-143.
Smith KK. 1986. Morphology and function of the tongue and hyoid apparatus in Varanus (Varanidae, Lacertilia). J Morphol. 187:261-287.
Smith KK, Kier WM. 1989. Trunks, tongues and tentacles: moving with skeletons of muscle. Am Sci. 77:28-35.
Smith KK, Mackay KA. 1990. The morphology of the intrinsic tongue musculature in snakes (Reptilia, Ophidia): functional and phylogenetic implications. J Morphol. 205:307-324.
Sokal RR, Rohlf FJ, editors. 1997. Biometry. The principles and practice of statistics in biological research. 3rd edition. New York: Freeman.
Stoddart DM. 1980. Olfaction in mammals. New York: Academic Press.
Toubeau G, Cotman C, Bels VL. 1994. Morphological and kinematic study of the tongue and buccal cavity in the lizard Anguis fragilis (Reptilia: Anguidae). Anat Rec. 240:423-433.
Ulinski PS. 1972. Tongue movements in the common boa (Constrictor constrictor). Anim Behav. 20:373-382.
Wyatt TD. 2003. Pheromones and animal behaviour: communication by smell and taste. Cambridge: Cambridge University Press.
